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aTyr Pharma, Inc. 2025-2026 Proxy Voting Records

Compiled from SEC Form N-PX filings. Page generated October 14, 2026.

7Reported items
103Asset managers
806Fund votes
May 11, 2026Meeting date

Results at a glance

No official results are on file for this meeting: no Form 8-K tally (Item 5.07) from aTyr Pharma, Inc. has been matched to it. The figures below are the funds’ own votes from Form N-PX. They are not the meeting result, so they are not used to judge dissent.

Material dissent: a director nominee under 80% of the votes cast, say-on-pay under 70%, any other item the board put forward under 80%, or a shareholder proposal with more than 50%, on the company’s own Form 8-K tally. How this is decided

Proxy season: 2025-2026

Explore aTyr Pharma, Inc. in the interactive database Compare manager voting policies

How asset managers voted at the aTyr Pharma, Inc. 2025-2026 meeting

Each item below shows how the 103 asset managers that disclosed a aTyr Pharma, Inc. vote in their SEC Form N-PX filings actually voted. Share totals are the shares those managers' funds voted, summed across every fund they report. These are reported items, not a count of the ballot: fund filings describe one ballot item in different words. Two descriptions are shown as one item only on evidence — the same text (or one cut short of the other), the same director nominee, or a match to the same row of aTyr Pharma, Inc.’s own Form 8-K tally — never because they merely read alike. A description that could not be joined that way is listed on its own, so one ballot item can appear more than once.

1. To elect three Class II directors, as nominated by the Company's Board of Directors (the "Board of…: Eric Benevich

No official resultsDIRECTOR ELECTIONS

Combines 4 wordings of this item as funds reported it.

99.9% of fund votes cast · no official result

FOR

Details: description as filed, votes by 35 asset managers

Description as filed by funds: To elect three Class II directors, as nominated by the Company's Board of Directors (the "Board of Directors"), to hold office until the 2029 annual meeting of stockholders or until their successors are duly elected and qualified: Eric Benevich

The 35 asset managers below reported 20,525,550 for, 8,686 withheld. The figure above is FOR ÷ (FOR + AGAINST + WITHHELD) of the shares these managers voted; abstained and unvoted shares are in neither half, and these managers are a subset of all shareholders, not the meeting result.

FOR: 20,525,550WITHHOLD: 8,686

Largest asset managers voting on “To elect three Class II directors, as nominated by the Company's Board of Directors (the "Board of Directors")” at aTyr Pharma, Inc., 2025-2026
Asset manager	For	Vote
Federated Hermes	8,220,788	For
Fidelity	7,759,704	For
Vanguard	2,033,412	For
BlackRock	1,606,882	For
Charles Schwab	208,534	For
TIAA	192,402	For
Equitable	102,174	For
Lincoln Financial	67,100	For
QUANTITATIVE MASTER SERIES LLC	60,500	For
Global X	44,089	For
AIG/SunAmerica	43,770	For
Goldman Sachs	41,449	For
Bridge Builder Trust	19,204	For
DWS	13,311	For
SEI	13,000	For
Nationwide	11,888	For
John Hancock	11,468	For
Blackstone Alternative Investment Funds	10,800	For
Principal	9,777	For
Voya	9,030	For
Calvert	8,881	For
T. Rowe Price	8,518	For
ProShares	8,313	For
Venerable Variable Insurance Trust	7,205	For
State Street	4,687	For

Showing the 25 largest of 35 asset managers. See all 35 in the interactive database.

2. To elect three Class II directors, as nominated by the Company's Board of Directors (the "Board…: Jane A. Gross, Ph.D.

No official resultsDIRECTOR ELECTIONS

Combines 4 wordings of this item as funds reported it.

99.1% of fund votes cast · no official result

FOR

Details: description as filed, votes by 35 asset managers

The 35 asset managers below reported 20,366,206 for, 168,029 withheld, 1 abstained. The figure above is FOR ÷ (FOR + AGAINST + WITHHELD) of the shares these managers voted; abstained and unvoted shares are in neither half, and these managers are a subset of all shareholders, not the meeting result.

FOR: 20,366,206ABSTAIN: 1WITHHOLD: 168,029

Largest asset managers voting on “To elect three Class II directors, as nominated by the Company's Board of Directors (the "Board of Directors")” at aTyr Pharma, Inc., 2025-2026
Asset manager	For	Abstain	Withheld	Vote
Federated Hermes	8,220,788	0	0	For
Fidelity	7,759,704	0	0	For
Vanguard	2,029,435	1	3,976	For
BlackRock	1,606,882	0	0	For
Charles Schwab	208,534	0	0	For
TIAA	192,402	0	0	For
Equitable	65,294	0	36,880	Mixed
Lincoln Financial	67,100	0	0	For
QUANTITATIVE MASTER SERIES LLC	60,500	0	0	For
Global X	44,089	0	0	For
AIG/SunAmerica	0	0	43,770	Withhold
Goldman Sachs	41,449	0	0	For
Bridge Builder Trust	19,204	0	0	For
DWS	0	0	13,311	Withhold
SEI	13,000	0	0	For
Nationwide	11,888	0	0	For
John Hancock	0	0	11,468	Withhold
Blackstone Alternative Investment Funds	0	0	10,800	Withhold
Principal	0	0	9,777	Withhold
Voya	0	0	9,030	Withhold
Calvert	0	0	8,881	Withhold
T. Rowe Price	8,518	0	0	For
ProShares	0	0	8,313	Withhold
Venerable Variable Insurance Trust	7,205	0	0	For
State Street	4,687	0	0	For

Showing the 25 largest of 35 asset managers. See all 35 in the interactive database.

3. To elect three Class II directors, as nominated by the Company's Board of Directors (the "Board…: Timothy P. Coughlin

No official resultsDIRECTOR ELECTIONS

Combines 4 wordings of this item as funds reported it.

99.9% of fund votes cast · no official result

FOR

Details: description as filed, votes by 35 asset managers

The 35 asset managers below reported 20,521,067 for, 13,169 withheld. The figure above is FOR ÷ (FOR + AGAINST + WITHHELD) of the shares these managers voted; abstained and unvoted shares are in neither half, and these managers are a subset of all shareholders, not the meeting result.

FOR: 20,521,067WITHHOLD: 13,169

Largest asset managers voting on “To elect three Class II directors, as nominated by the Company's Board of Directors (the "Board of Directors")” at aTyr Pharma, Inc., 2025-2026
Asset manager	For	Withheld	Vote
Federated Hermes	8,220,788	0	For
Fidelity	7,759,704	0	For
Vanguard	2,029,436	3,976	For
BlackRock	1,606,882	0	For
Charles Schwab	208,534	0	For
TIAA	192,402	0	For
Equitable	102,174	0	For
Lincoln Financial	67,100	0	For
QUANTITATIVE MASTER SERIES LLC	60,500	0	For
Global X	44,089	0	For
AIG/SunAmerica	43,770	0	For
Goldman Sachs	41,449	0	For
Bridge Builder Trust	19,204	0	For
DWS	13,311	0	For
SEI	13,000	0	For
Nationwide	11,888	0	For
John Hancock	11,468	0	For
Blackstone Alternative Investment Funds	10,800	0	For
Principal	9,777	0	For
Voya	9,030	0	For
Calvert	8,881	0	For
T. Rowe Price	8,518	0	For
ProShares	8,313	0	For
Venerable Variable Insurance Trust	7,205	0	For
State Street	4,687	0	For

Showing the 25 largest of 35 asset managers. See all 35 in the interactive database.

4. To approve an amendment to the Company's Restated Certificate of Incorporation to increase the number of authorized…

No official resultsCAPITAL STRUCTURE

Combines 4 wordings of this item as funds reported it.

99.9% of fund votes cast · no official result

FOR

Details: description as filed, votes by 35 asset managers

Description as filed by funds: To approve an amendment to the Company's Restated Certificate of Incorporation to increase the number of authorized shares of common stock from 170,000,000 to 340,000,000 shares; and

The 35 asset managers below reported 20,531,009 for, 3,227 against. The figure above is FOR ÷ (FOR + AGAINST + WITHHELD) of the shares these managers voted; abstained and unvoted shares are in neither half, and these managers are a subset of all shareholders, not the meeting result.

FOR: 20,531,009AGAINST: 3,227

Largest asset managers voting on “To approve an amendment to the Company's Restated Certificate of Incorporation to increase the number of autho” at aTyr Pharma, Inc., 2025-2026
Asset manager	For	Against	Vote
Federated Hermes	8,220,788	0	For
Fidelity	7,759,704	0	For
Vanguard	2,030,692	2,720	For
BlackRock	1,606,882	0	For
Charles Schwab	208,534	0	For
TIAA	192,402	0	For
Equitable	102,174	0	For
Lincoln Financial	67,100	0	For
QUANTITATIVE MASTER SERIES LLC	60,500	0	For
Global X	44,089	0	For
AIG/SunAmerica	43,770	0	For
Goldman Sachs	41,449	0	For
Bridge Builder Trust	19,204	0	For
DWS	13,311	0	For
SEI	13,000	0	For
Nationwide	11,888	0	For
John Hancock	11,468	0	For
Blackstone Alternative Investment Funds	10,800	0	For
Principal	9,777	0	For
Voya	9,030	0	For
Calvert	8,881	0	For
T. Rowe Price	8,518	0	For
ProShares	8,313	0	For
Venerable Variable Insurance Trust	7,205	0	For
State Street	4,687	0	For

Showing the 25 largest of 35 asset managers. See all 35 in the interactive database.

5. To approve an amendment to the aTyr Pharma, Inc. 2015 Stock Option and Incentive Plan, as amended;

No official resultsCOMPENSATION

Combines 4 wordings of this item as funds reported it.

98% of fund votes cast · no official result

FOR

Details: votes by 35 asset managers

The 35 asset managers below reported 20,215,434 for, 318,802 against. The figure above is FOR ÷ (FOR + AGAINST + WITHHELD) of the shares these managers voted; abstained and unvoted shares are in neither half, and these managers are a subset of all shareholders, not the meeting result.

FOR: 20,215,434AGAINST: 318,802

Largest asset managers voting on “To approve an amendment to the aTyr Pharma, Inc. 2015 Stock Option and Incentive Plan, as amended;” at aTyr Pharma, Inc., 2025-2026
Asset manager	For	Against	Vote
Federated Hermes	8,220,788	0	For
Fidelity	7,759,704	0	For
Vanguard	2,033,410	2	For
BlackRock	1,606,882	0	For
Charles Schwab	208,534	0	For
TIAA	0	192,402	Against
Equitable	102,174	0	For
Lincoln Financial	67,100	0	For
QUANTITATIVE MASTER SERIES LLC	60,500	0	For
Global X	44,089	0	For
AIG/SunAmerica	0	43,770	Against
Goldman Sachs	41,449	0	For
Bridge Builder Trust	19,204	0	For
DWS	0	13,311	Against
SEI	13,000	0	For
Nationwide	11,888	0	For
John Hancock	0	11,468	Against
Blackstone Alternative Investment Funds	0	10,800	Against
Principal	0	9,777	Against
Voya	0	9,030	Against
Calvert	8,881	0	For
T. Rowe Price	0	8,518	Against
ProShares	0	8,313	Against
Venerable Variable Insurance Trust	7,205	0	For
State Street	4,687	0	For

Showing the 25 largest of 35 asset managers. See all 35 in the interactive database.

6. To approve, on an advisory basis, the compensation of the Company's named executive officers;

No official results Fund totals may overlapSECTION 14A SAY-ON-PAY VOTES

Combines 3 wordings of this item as funds reported it.

99.6% Fund and manager reports combined; may count the same shares twice · no official result

FOR

Details: votes by 103 asset managers

Fund voting reports and institutional-manager reports both cover this item. An institutional manager can report shares that a fund it advises also reports, so the two are shown separately below and never added together, and their sum is not shown (fund and manager reports combined may count the same shares twice). The figure above is FOR ÷ (FOR + AGAINST + WITHHELD) of the shares these managers voted; abstained and unvoted shares are in neither half, and these managers are a subset of all shareholders, not the meeting result.

Fund voting reports: 35 asset managers voting on “To approve, on an advisory basis, the compensation of the Company's named executive officers;” at aTyr Pharma, Inc., 2025-2026
Asset manager	For	Against	Vote
Federated Hermes	8,220,788	0	For
Fidelity	7,759,704	0	For
Vanguard	2,033,411	1	For
BlackRock	1,606,882	0	For
Charles Schwab	208,534	0	For
TIAA	192,402	0	For
Equitable	102,174	0	For
Lincoln Financial	67,100	0	For
QUANTITATIVE MASTER SERIES LLC	60,500	0	For
Global X	44,089	0	For
AIG/SunAmerica	43,770	0	For
Goldman Sachs	41,449	0	For
Bridge Builder Trust	19,204	0	For
DWS	13,311	0	For
SEI	13,000	0	For
Nationwide	11,888	0	For
John Hancock	11,468	0	For
Blackstone Alternative Investment Funds	10,800	0	For
Principal	9,777	0	For
Voya	9,030	0	For
Calvert	8,881	0	For
T. Rowe Price	8,518	0	For
ProShares	8,313	0	For
Venerable Variable Insurance Trust	7,205	0	For
State Street	4,687	0	For

Fund voting reports, all 35 managers: 20,525,042 for, 9,194 against. Showing the 25 largest; see all in the interactive database.

Institutional-manager reports: 81 asset managers voting on “To approve, on an advisory basis, the compensation of the Company's named executive officers;” at aTyr Pharma, Inc., 2025-2026
Asset manager	For	Against	Vote
Fidelity	12,186,117	0	For
Federated Hermes	8,220,788	0	For
GEODE CAPITAL MANAGEMENT, LLC	2,040,035	0	For
State Street	911,080	3,071	For
Wilkinson Global Asset Management LLC	819,550	0	For
BlackRock	671,193	232	For
Northern Trust	347,067	2,812	For
MELLON INVESTMENTS Corp	230,953	0	For
Nuveen	226,231	0	For
Charles Schwab	210,737	0	For
AQR	132,019	0	For
Vanguard	121,787	0	For
Goldman Sachs	97,945	0	For
MARSHALL WACE, LLP	89,103	0	For
Russell Investments	88,520	0	For
Dimensional	0	83,138	Against
AllianceBernstein	64,480	0	For
BANK OF AMERICA NA	46,536	0	For
Invesco	0	43,219	Against
T. Rowe Price	41,260	0	For
Two Sigma	34,456	0	For
NEW YORK STATE COMMON RETIREMENT FUND	27,500	0	For
Citadel	25,091	0	For
STIFEL NICOLAUS & CO INC \MO\	23,200	0	For
Neuberger Berman	19,884	0	For

Institutional-manager reports, all 81 managers: 26,895,500 for, 170,252 against. Showing the 25 largest; see all in the interactive database.

7. To ratify the appointment of Ernst & Young LLP as the independent registered public accounting firm of the Company for…

No official resultsAUDIT-RELATED

Combines 2 wordings of this item as funds reported it.

100% of fund votes cast · no official result

FOR

Details: description as filed, votes by 35 asset managers

Description as filed by funds: To ratify the appointment of Ernst & Young LLP as the independent registered public accounting firm of the Company for its fiscal year ending December 31, 2026;

The 35 asset managers below reported 20,534,236 for. The figure above is FOR ÷ (FOR + AGAINST + WITHHELD) of the shares these managers voted; abstained and unvoted shares are in neither half, and these managers are a subset of all shareholders, not the meeting result.

FOR: 20,534,236

Largest asset managers voting on “To ratify the appointment of Ernst & Young LLP as the independent registered public accounting firm of the Com” at aTyr Pharma, Inc., 2025-2026
Asset manager	For	Vote
Federated Hermes	8,220,788	For
Fidelity	7,759,704	For
Vanguard	2,033,412	For
BlackRock	1,606,882	For
Charles Schwab	208,534	For
TIAA	192,402	For
Equitable	102,174	For
Lincoln Financial	67,100	For
QUANTITATIVE MASTER SERIES LLC	60,500	For
Global X	44,089	For
AIG/SunAmerica	43,770	For
Goldman Sachs	41,449	For
Bridge Builder Trust	19,204	For
DWS	13,311	For
SEI	13,000	For
Nationwide	11,888	For
John Hancock	11,468	For
Blackstone Alternative Investment Funds	10,800	For
Principal	9,777	For
Voya	9,030	For
Calvert	8,881	For
T. Rowe Price	8,518	For
ProShares	8,313	For
Venerable Variable Insurance Trust	7,205	For
State Street	4,687	For

Showing the 25 largest of 35 asset managers. See all 35 in the interactive database.

Largest aTyr Pharma, Inc. shareholders voting in 2025-2026

Ranked separately for each type of Form N-PX report. An institutional manager can report shares that a fund it advises also reports, so the two tables can list the same shares and are never added together. Each is ranked by the shares voted on the item that the most managers of that type reported, shown as a share of the 98,051,212 shares outstanding at the time of that meeting.

Fund voting reports: top aTyr Pharma, Inc. shareholders by shares voted on “To approve, on an advisory basis, the compensation of the Company's named executive officers;”, 2025-2026
#	Asset manager	% of shares outstanding
1	Federated Hermes	8.38%
2	Fidelity	7.91%
3	Vanguard	2.07%
4	BlackRock	1.64%
5	Charles Schwab	0.21%
6	TIAA	0.20%
7	Equitable	0.10%
8	Lincoln Financial	0.07%
9	QUANTITATIVE MASTER SERIES LLC	0.06%
10	Global X	0.04%

Institutional-manager reports: top aTyr Pharma, Inc. shareholders by shares voted on “To approve, on an advisory basis, the compensation of the Company's named executive officers;”, 2025-2026
#	Asset manager	% of shares outstanding
1	Fidelity	12.43%
2	Federated Hermes	8.38%
3	GEODE CAPITAL MANAGEMENT, LLC	2.08%
4	State Street	0.93%
5	Wilkinson Global Asset Management LLC	0.84%
6	BlackRock	0.68%
7	Northern Trust	0.36%
8	MELLON INVESTMENTS Corp	0.24%
9	Nuveen	0.23%
10	Charles Schwab	0.21%

Reported aTyr Pharma, Inc. holdings and ownership

As disclosed to the SEC in Form 13F (quarter-end holdings of 30 large managers), Schedules 13D and 13G (beneficial ownership above 5%) and the company’s own proxy statement (DEF 14A). Each row names its form.

aTyr Pharma, Inc. holdings and ownership disclosed for the 2025-2026 proxy season
Holder	% outstanding	Disclosure
FMR (Fidelity)	15.00%	13F
1001 Liberty Avenue	8.68%	DEF14A
Federated Hermes, Inc.	8.68%	13G
BlackRock	6.57%	13F
Vanguard Group	5.06%	13F
Geode Capital	2.43%	13F
State Street	1.79%	13F
Millennium Management	1.67%	13F
Morgan Stanley	1.43%	13F
Two Sigma	1.17%	13F

Percentages above are of 98,051,212 shares outstanding, as reported by aTyr Pharma, Inc. on its Form 10-Q dated May 11, 2026 (see the filing on EDGAR). This is the count contemporaneous with the 2025-2026 meeting, not today’s. Each holding is as of its own filing, which may be dated before or after the meeting.

About this page

Every figure above is derived from filings made with the US Securities and Exchange Commission: Form N-PX, in which registered funds disclose how they voted every proxy and institutional managers disclose their votes on executive pay (say-on-pay, say-on-frequency and golden parachutes), and Form 8-K Item 5.07, in which the company reports its own meeting results. Shares outstanding are taken from aTyr Pharma, Inc.’s 10-Q dated May 11, 2026. This page is a static snapshot rebuilt weekly on October 14, 2026; a live search always shows the current data.

At aTyr Pharma, Inc.'s shareholder meeting held May 11, 2026, in the 2025-2026 proxy season, 103 asset managers reported how they voted in their SEC Form N-PX filings, covering 806 separate fund positions. Their filings are grouped here into 7 reported items; because filers word the same ballot item differently, that can be more than the number of items on the ballot. No official Form 8-K vote tally has been matched to this meeting, so this page states no certified outcome for its most widely held ballot item. On that item — To approve, on an advisory basis, the compensation of the Company's named executive officers; — the reporting funds and managers cast 99.6% of the shares they voted in favour (FOR divided by FOR plus AGAINST plus WITHHELD; abstained and unvoted shares are excluded because neither took a side). Fund and manager reports are combined in it and may count the same shares twice: it adds fund reports to institutional-manager reports, which can cover the same shares. It is not the company's certified result.

aTyr Pharma, Inc. proxy season coverage: 2025-2026 (this page).

Download data (CSV): ballot items · votes by manager · column guide.